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A Health Upgrade permanently increases the player's maximum Health capacity.
Health Upgrades can be purchased at Gatherer's Gardens throughout the game for a sum of 80 ADAM, and should be the first thing that new players purchase (provided they choose to save the Little Sisters). They are also a reward for Little Sister research.
Health Upgrades return as a purchasable upgrade at Gatherer's Gardens, although the price has now been reduced to 40 ADAM.